:root {
  --paper: #f1ece3;
  --paper-light: #faf7f1;
  --ink: #211c1b;
  --muted: #665f5b;
  --plum: #651f3a;
  --plum-dark: #35101f;
  --plum-soft: #a66a7e;
  --ice: #b9d1df;
  --ice-light: #dce9ef;
  --moss: #626f65;
  --line: rgba(33, 28, 27, 0.18);
  --display: Georgia, "Times New Roman", serif;
  --sans: "Helvetica Neue", Helvetica, Arial, sans-serif;
  --max: 1440px;
  --pad: clamp(1.25rem, 3.5vw, 4.5rem);
  --radius-lg: 0 6rem 0 6rem;
  --shadow: 0 1.5rem 4rem rgba(37, 20, 26, 0.14);
}

* {
  box-sizing: border-box;
}

html {
  scroll-behavior: smooth;
}

body {
  margin: 0;
  background: var(--paper);
  color: var(--ink);
  font-family: var(--sans);
  font-size: 16px;
  line-height: 1.55;
  -webkit-font-smoothing: antialiased;
}

body.menu-open {
  overflow: hidden;
}

::selection {
  background: var(--plum);
  color: white;
}

img {
  display: block;
  max-width: 100%;
}

a {
  color: inherit;
  text-decoration: none;
}

button,
input {
  font: inherit;
}

button {
  color: inherit;
}

p,
h1,
h2,
h3,
h4 {
  margin-top: 0;
}

.skip-link {
  position: fixed;
  z-index: 1000;
  top: 0.75rem;
  left: 0.75rem;
  padding: 0.7rem 1rem;
  background: white;
  color: var(--ink);
  transform: translateY(-150%);
  transition: transform 180ms ease;
}

.skip-link:focus {
  transform: translateY(0);
}

.site-header {
  position: sticky;
  z-index: 100;
  top: 0;
  background: rgba(241, 236, 227, 0.93);
  border-bottom: 1px solid transparent;
  backdrop-filter: blur(15px);
  transition: border-color 200ms ease, background 200ms ease;
}

.site-header.is-scrolled {
  border-color: var(--line);
}

.nav-shell {
  width: min(100%, var(--max));
  min-height: 5.8rem;
  margin: 0 auto;
  padding: 0 var(--pad);
  display: flex;
  align-items: center;
  justify-content: space-between;
  gap: 2rem;
}

.brand {
  display: inline-flex;
  align-items: center;
  gap: 0.8rem;
  min-width: 13rem;
  max-width: 27rem;
  padding-block: .75rem;
  font-size: 0.74rem;
  font-weight: 700;
  line-height: 1.18;
  letter-spacing: 0.12em;
  text-transform: uppercase;
}

.brand-mark {
  display: block;
  width: 2.5rem;
  height: 2.5rem;
  flex: 0 0 auto;
}

.nav-links {
  display: flex;
  align-items: center;
  gap: clamp(1.3rem, 2.6vw, 3.2rem);
}

.nav-link {
  position: relative;
  padding: 0.6rem 0;
  font-size: 0.77rem;
  font-weight: 700;
  letter-spacing: 0.1em;
  text-transform: uppercase;
}

.nav-link::after {
  content: "";
  position: absolute;
  right: 0;
  bottom: 0.25rem;
  left: 0;
  height: 1px;
  background: currentColor;
  transform: scaleX(0);
  transform-origin: right;
  transition: transform 250ms ease;
}

.nav-link:hover::after,
.nav-link.active::after {
  transform: scaleX(1);
  transform-origin: left;
}

.nav-link.active {
  color: var(--plum);
}

.menu-toggle {
  display: none;
  width: 3rem;
  height: 3rem;
  padding: 0;
  border: 1px solid var(--line);
  border-radius: 50%;
  background: transparent;
  cursor: pointer;
}

.menu-toggle span,
.menu-toggle::before,
.menu-toggle::after {
  content: "";
  display: block;
  width: 1.6rem;
  height: 1px;
  margin: 0.34rem auto;
  background: currentColor;
  transition: transform 200ms ease, opacity 200ms ease;
}

.menu-open .menu-toggle span {
  opacity: 0;
}

.menu-open .menu-toggle::before {
  transform: translateY(0.43rem) rotate(45deg);
}

.menu-open .menu-toggle::after {
  transform: translateY(-0.43rem) rotate(-45deg);
}

main {
  overflow: hidden;
}

.shell {
  width: min(100%, var(--max));
  margin: 0 auto;
  padding-right: var(--pad);
  padding-left: var(--pad);
}

.section {
  padding-top: clamp(5.5rem, 10vw, 10rem);
  padding-bottom: clamp(5.5rem, 10vw, 10rem);
}

.section-tight {
  padding-top: clamp(3.5rem, 6vw, 6rem);
  padding-bottom: clamp(3.5rem, 6vw, 6rem);
}

.eyebrow {
  display: flex;
  align-items: center;
  gap: 0.65rem;
  margin-bottom: 1.4rem;
  color: var(--plum);
  font-size: 0.72rem;
  font-weight: 700;
  letter-spacing: 0.16em;
  text-transform: uppercase;
}

.eyebrow::before {
  content: "";
  width: 2.25rem;
  height: 1px;
  background: currentColor;
}

.eyebrow.light {
  color: var(--ice-light);
}

.display-xl,
.display-lg,
.display-md {
  font-family: var(--display);
  font-weight: 400;
  letter-spacing: -0.045em;
  line-height: 0.92;
}

.display-xl {
  font-size: clamp(4.2rem, 9.8vw, 9.6rem);
}

.display-lg {
  font-size: clamp(3.3rem, 7vw, 7rem);
}

.display-md {
  font-size: clamp(2.5rem, 4.8vw, 4.9rem);
}

.serif-italic {
  font-style: italic;
}

.lead {
  max-width: 42rem;
  color: var(--muted);
  font-family: var(--display);
  font-size: clamp(1.3rem, 2vw, 1.75rem);
  line-height: 1.42;
}

.body-lg {
  color: var(--muted);
  font-size: clamp(1.05rem, 1.5vw, 1.25rem);
  line-height: 1.7;
}

.button,
.text-link {
  display: inline-flex;
  align-items: center;
  justify-content: center;
  gap: 1rem;
  font-size: 0.73rem;
  font-weight: 700;
  letter-spacing: 0.13em;
  text-transform: uppercase;
}

.button {
  min-height: 3.5rem;
  padding: 0.9rem 1.35rem;
  border: 1px solid var(--ink);
  background: transparent;
  transition: color 200ms ease, background 200ms ease, transform 200ms ease;
}

.button:hover {
  background: var(--ink);
  color: white;
  transform: translateY(-2px);
}

.button.light {
  border-color: rgba(255,255,255,0.65);
  color: white;
}

.button.light:hover {
  background: white;
  color: var(--ink);
}

.arrow {
  font-size: 1.2rem;
  font-weight: 400;
  transition: transform 200ms ease;
}

.button:hover .arrow,
.text-link:hover .arrow {
  transform: translateX(0.3rem);
}

.text-link {
  padding-bottom: 0.35rem;
  border-bottom: 1px solid currentColor;
}

.home-hero {
  position: relative;
  min-height: calc(100vh - 5.8rem);
  min-height: calc(100svh - 5.8rem);
  padding-top: clamp(3rem, 5vw, 5rem);
  padding-bottom: 3rem;
}

.home-hero-grid {
  display: grid;
  grid-template-columns: minmax(0, 1.22fr) minmax(23rem, 0.78fr);
  grid-template-rows: auto auto;
  align-items: end;
  gap: 0 clamp(2rem, 5vw, 6rem);
}

.home-hero-copy {
  position: relative;
  z-index: 2;
  padding-top: clamp(2rem, 7vw, 7rem);
}

.home-hero-copy h1 {
  max-width: 9.4em;
  margin-bottom: 2.4rem;
}

.home-research-title {
  font-family: var(--display);
  font-size: clamp(2.25rem, 4.3vw, 4.5rem);
  font-weight: 400;
  letter-spacing: -0.035em;
  line-height: 1.12;
}

.home-hero-copy .hero-line {
  display: block;
}

.home-hero-copy .hero-line:nth-child(2) {
  color: var(--plum);
  transform: translateX(clamp(0rem, 6vw, 6rem));
}

.home-hero-image-wrap {
  position: relative;
  align-self: stretch;
  min-height: 39rem;
  overflow: hidden;
  border-radius: var(--radius-lg);
  box-shadow: var(--shadow);
}

.home-hero-image {
  position: absolute;
  inset: 0;
  width: 100%;
  height: 100%;
  object-fit: cover;
  object-position: 62% center;
  transform: scale(1.02);
}

.image-label {
  position: absolute;
  right: 1.1rem;
  bottom: 1.1rem;
  max-width: 11rem;
  padding: 0.7rem 0.85rem;
  background: rgba(250, 247, 241, 0.92);
  font-size: 0.65rem;
  font-weight: 700;
  letter-spacing: 0.11em;
  line-height: 1.3;
  text-transform: uppercase;
  backdrop-filter: blur(8px);
}

.hero-intro {
  display: grid;
  grid-template-columns: 1fr;
  gap: clamp(1.5rem, 4vw, 5rem);
  align-items: end;
  padding-top: 2.5rem;
}

.hero-intro p {
  max-width: 40rem;
  margin-bottom: 0;
}

.mission-grid {
  display: grid;
  grid-template-columns: minmax(0, 0.85fr) minmax(24rem, 1.15fr);
  gap: clamp(3rem, 8vw, 9rem);
  align-items: center;
}

.transplant-network {
  margin: 0;
  min-width: 0;
}

.network-map {
  display: block;
  width: 100%;
  height: auto;
  overflow: visible;
}

.network-nodes circle {
  fill: var(--plum);
  stroke: var(--paper);
  stroke-width: 2;
}

.network-traveler {
  fill: var(--plum);
  stroke: var(--paper-light);
  stroke-width: 1;
  stroke-linejoin: round;
  pointer-events: none;
}

.transplant-network figcaption {
  display: flex;
  flex-wrap: wrap;
  justify-content: space-between;
  gap: .75rem;
  margin-top: 1rem;
  color: var(--muted);
  font-size: .7rem;
  line-height: 1.5;
}

.network-toggle {
  padding: 0;
  border: 0;
  background: none;
  color: inherit;
  font: inherit;
  text-decoration: underline;
  text-underline-offset: .2em;
  cursor: pointer;
}

@media (prefers-reduced-motion: reduce) {
  .network-traveler, .network-toggle { display: none; }
}

.funding-project-title {
  margin: 1.5rem 0;
  font-size: 1.35rem;
  line-height: 1.4;
}

#funding {
  scroll-margin-top: 6rem;
}

.team-section {
  min-height: 75svh;
  display: flex;
  flex-direction: column;
  justify-content: center;
}

.team-section .section-heading-row {
  margin-bottom: 0;
}

.funding-section {
  min-height: 90svh;
  display: flex;
  align-items: center;
  background: var(--paper-light);
  border-top: 1px solid var(--line);
}

.vertical-note {
  position: absolute;
  right: 0;
  bottom: 0;
  padding: 0.75rem 0;
  color: var(--muted);
  font-size: 0.67rem;
  font-weight: 700;
  letter-spacing: 0.12em;
  text-transform: uppercase;
  writing-mode: vertical-rl;
  transform: rotate(180deg);
}

.mission-copy h2 {
  margin-bottom: 2.5rem;
}

.mission-copy .body-lg {
  max-width: 37rem;
}

.research-metrics {
  display: flex;
  flex-wrap: wrap;
  gap: 1.5rem 3rem;
  margin-top: 2rem;
}

.research-metrics > div {
  display: flex;
  flex-direction: column;
  gap: .4rem;
}

.research-metrics strong {
  color: var(--plum);
  font-family: var(--display);
  font-size: 2.4rem;
  font-weight: 400;
  line-height: 1.1;
}

.research-metrics-note {
  color: var(--muted);
  font-size: .75rem;
  line-height: 1.6;
  margin-top: 1rem;
}

.stat-row {
  display: grid;
  grid-template-columns: repeat(3, 1fr);
  gap: 1px;
  margin-top: 3.5rem;
  background: var(--line);
  border: 1px solid var(--line);
}

.stat {
  min-height: 9rem;
  padding: 1.3rem;
  background: var(--paper);
}

.stat strong {
  display: block;
  margin-bottom: 1.2rem;
  color: var(--plum);
  font-family: var(--display);
  font-size: clamp(1.8rem, 3.4vw, 3rem);
  font-weight: 400;
  line-height: 1;
}

.stat span {
  display: block;
  color: var(--muted);
  font-size: 0.69rem;
  font-weight: 700;
  letter-spacing: 0.09em;
  line-height: 1.4;
  text-transform: uppercase;
}

.dark-section {
  background: var(--plum-dark);
  color: white;
}

.dark-section .body-lg,
.dark-section .lead {
  color: rgba(255,255,255,0.68);
}

.section-heading-row {
  display: flex;
  align-items: end;
  justify-content: space-between;
  gap: 2rem;
  margin-bottom: clamp(3rem, 6vw, 6rem);
}

.section-heading-row h2 {
  max-width: 10em;
  margin-bottom: 0;
}

.pillar-grid {
  display: grid;
  grid-template-columns: repeat(4, minmax(0, 1fr));
  border-top: 1px solid rgba(255,255,255,0.2);
}

.pillar {
  min-height: 27rem;
  padding: 2rem clamp(1.3rem, 3vw, 3.5rem) 2.5rem 0;
  border-right: 1px solid rgba(255,255,255,0.2);
}

.pillar + .pillar {
  padding-left: clamp(1.3rem, 3vw, 3.5rem);
}

.pillar:last-child {
  border-right: 0;
}

.pillar-number {
  display: block;
  margin-bottom: clamp(5rem, 9vw, 9rem);
  color: var(--ice);
  font-family: var(--display);
  font-size: 1.2rem;
}

.pillar h3 {
  margin-bottom: 1.3rem;
  font-family: var(--display);
  font-size: clamp(1.65rem, 2.6vw, 2.35rem);
  font-weight: 400;
  line-height: 1.12;
}

.pillar h3 a:hover,
.pillar h3 a:focus-visible {
  text-decoration: underline;
  text-underline-offset: .15em;
}

.pillar p {
  margin-bottom: 0;
  color: rgba(255,255,255,0.62);
}

.grant-band {
  display: grid;
  grid-template-columns: 1.25fr 0.75fr;
  gap: clamp(3rem, 8vw, 8rem);
  align-items: start;
}

.grant-band h2 {
  margin-bottom: 2rem;
}

.grant-facts {
  margin: 0;
  border-top: 1px solid var(--line);
}

.grant-fact {
  display: grid;
  grid-template-columns: 8.5rem 1fr;
  gap: 1rem;
  padding: 1.15rem 0;
  border-bottom: 1px solid var(--line);
}

.grant-fact dt {
  color: var(--muted);
  font-size: 0.67rem;
  font-weight: 700;
  letter-spacing: 0.1em;
  text-transform: uppercase;
}

.grant-fact dd {
  margin: 0;
}

.page-hero {
  padding-top: clamp(5rem, 9vw, 9rem);
  padding-bottom: clamp(4.5rem, 8vw, 8rem);
}

.page-hero-grid {
  display: grid;
  grid-template-columns: minmax(0, 1.3fr) minmax(20rem, 0.7fr);
  gap: clamp(3rem, 8vw, 9rem);
  align-items: end;
}

.page-hero h1 {
  max-width: 10em;
  margin-bottom: 0;
}

.page-hero-aside {
  padding-bottom: 0.5rem;
}

.page-hero-aside .lead {
  margin-bottom: 2.2rem;
}

.team-lead {
  display: grid;
  grid-template-columns: repeat(3, 1fr);
  gap: 1rem;
}

.pi-card {
  position: relative;
  display: flex;
  flex-direction: column;
  min-height: 31rem;
  padding: clamp(1.5rem, 3vw, 2.5rem);
  overflow: hidden;
  background: var(--plum-dark);
  color: white;
}

.pi-card:nth-child(2) {
  background: var(--ice);
  color: var(--ink);
}

.pi-card:nth-child(3) {
  background: var(--plum);
}

.pi-role {
  position: relative;
  z-index: 2;
  color: inherit;
  font-size: 0.67rem;
  font-weight: 700;
  letter-spacing: 0.13em;
  text-transform: uppercase;
  opacity: 0.72;
}

.pi-monogram {
  position: absolute;
  top: 52%;
  left: 50%;
  color: rgba(255,255,255,0.09);
  font-family: var(--display);
  font-size: clamp(10rem, 19vw, 18rem);
  letter-spacing: -0.1em;
  line-height: 0.7;
  transform: translate(-52%, -50%);
}

.pi-card:nth-child(2) .pi-monogram {
  color: rgba(33,28,27,0.08);
}

.pi-card-content {
  position: relative;
  z-index: 2;
  margin-top: auto;
}

.pi-card h2 {
  margin-bottom: 0.8rem;
  font-family: var(--display);
  font-size: clamp(2.2rem, 3.8vw, 3.6rem);
  font-weight: 400;
  line-height: 0.98;
}

.pi-card p {
  max-width: 20rem;
  margin-bottom: 0;
  font-size: 0.88rem;
  opacity: 0.68;
}

.team-grid {
  display: grid;
  grid-template-columns: repeat(2, 1fr);
  border-top: 1px solid var(--line);
}

.team-member {
  display: grid;
  grid-template-columns: minmax(0, 1fr);
  gap: clamp(1.4rem, 3vw, 3rem);
  min-height: 17rem;
  padding: 2.5rem 2.5rem 2.5rem 0;
  border-bottom: 1px solid var(--line);
}

.team-member:nth-child(even) {
  padding-left: 2.5rem;
  border-left: 1px solid var(--line);
}

.team-number {
  color: var(--plum);
  font-family: var(--display);
  font-size: 1.1rem;
}

.team-member h3 {
  margin-bottom: 0.35rem;
  font-family: var(--display);
  font-size: clamp(1.8rem, 3vw, 2.8rem);
  font-weight: 400;
  line-height: 1;
}

.team-title {
  margin-bottom: 1.5rem;
  color: var(--plum);
  font-size: 0.68rem;
  font-weight: 700;
  letter-spacing: 0.1em;
  text-transform: uppercase;
}

.team-bio {
  max-width: 26rem;
  margin-bottom: 0;
  color: var(--muted);
}

.mock-note {
  margin-top: 2rem;
  color: var(--muted);
  font-size: 0.76rem;
}

.disciplines {
  display: flex;
  flex-wrap: wrap;
  gap: 0.7rem;
  margin-top: 1.6rem;
}

.tag {
  padding: 0.45rem 0.7rem;
  border: 1px solid var(--line);
  border-radius: 999px;
  color: var(--muted);
  font-size: 0.66rem;
  font-weight: 700;
  letter-spacing: 0.08em;
  text-transform: uppercase;
}

.pub-toolbar {
  display: flex;
  flex-wrap: wrap;
  gap: 0.7rem;
  margin-bottom: 2.5rem;
}

.filter-button {
  padding: 0.7rem 1rem;
  border: 1px solid var(--line);
  border-radius: 99px;
  background: transparent;
  color: var(--muted);
  font-size: 0.68rem;
  font-weight: 700;
  letter-spacing: 0.08em;
  text-transform: uppercase;
  cursor: pointer;
  transition: background 180ms ease, color 180ms ease;
}

.filter-button.active,
.filter-button:hover {
  background: var(--ink);
  color: white;
}

.publication-list {
  border-top: 1px solid var(--ink);
}

.publication {
  display: grid;
  grid-template-columns: 6rem minmax(0, 1fr) 2.5rem;
  gap: clamp(1.2rem, 3vw, 3rem);
  align-items: start;
  padding: clamp(1.8rem, 3vw, 3rem) 0;
  border-bottom: 1px solid var(--line);
  transition: opacity 220ms ease, transform 220ms ease;
}

.publication.is-hidden {
  display: none;
}

.pub-year {
  color: var(--plum);
  font-family: var(--display);
  font-size: 1.4rem;
}

.publication h2 {
  max-width: 40em;
  margin-bottom: 1rem;
  font-family: var(--display);
  font-size: clamp(1.5rem, 2.5vw, 2.35rem);
  font-weight: 400;
  line-height: 1.22;
}

.pub-meta,
.pub-authors {
  margin-bottom: 0.4rem;
  color: var(--muted);
  font-size: 0.82rem;
}

.pub-type {
  display: inline-block;
  margin-top: 0.9rem;
  color: var(--plum);
  font-size: 0.64rem;
  font-weight: 700;
  letter-spacing: 0.1em;
  text-transform: uppercase;
}

.pub-arrow {
  display: grid;
  width: 2.3rem;
  height: 2.3rem;
  place-items: center;
  border: 1px solid var(--line);
  border-radius: 50%;
  transition: background 180ms ease, color 180ms ease, transform 180ms ease;
}

.publication:hover .pub-arrow {
  background: var(--plum);
  color: white;
  transform: translate(0.2rem, -0.2rem);
}

.citation-note {
  margin-top: 2rem;
  color: var(--muted);
  font-size: 0.78rem;
}

.comet-hero {
  position: relative;
  min-height: calc(100vh - 5.8rem);
  min-height: calc(100svh - 5.8rem);
  display: grid;
  align-items: end;
  overflow: hidden;
  background: #080a0c;
  color: white;
}

.comet-hero-image {
  position: absolute;
  inset: 0;
  width: 100%;
  height: 100%;
  object-fit: cover;
  object-position: center;
  opacity: 0.82;
}

.comet-hero::after {
  content: "";
  position: absolute;
  inset: 0;
  background: linear-gradient(90deg, rgba(5,7,8,0.75) 0%, rgba(5,7,8,0.12) 60%, rgba(5,7,8,0.3) 100%), linear-gradient(0deg, rgba(5,7,8,0.9), transparent 60%);
}

.comet-hero-content {
  position: relative;
  z-index: 2;
  padding-top: 7rem;
  padding-bottom: clamp(3rem, 6vw, 6rem);
}

.comet-hero h1 {
  max-width: 7.5em;
  margin-bottom: 1.6rem;
}

.comet-hero .lead {
  max-width: 34rem;
  margin-bottom: 2.3rem;
  color: rgba(255,255,255,0.75);
}

.comet-kicker {
  color: var(--ice);
  font-size: 0.7rem;
  font-weight: 700;
  letter-spacing: 0.15em;
  text-transform: uppercase;
}

.model-intro {
  display: grid;
  grid-template-columns: 0.75fr 1.25fr;
  gap: clamp(3rem, 9vw, 10rem);
  align-items: start;
}

.model-intro h2 {
  position: sticky;
  top: 8rem;
}

.model-intro-copy .lead {
  margin-bottom: 2rem;
}

.process {
  margin-top: 4rem;
  border-top: 1px solid var(--line);
}

.process-step {
  display: grid;
  grid-template-columns: 4rem 1fr;
  gap: 1.2rem;
  padding: 1.7rem 0;
  border-bottom: 1px solid var(--line);
}

.process-step span {
  color: var(--plum);
  font-family: var(--display);
  font-size: 1.2rem;
}

.process-step h3 {
  margin-bottom: 0.4rem;
  font-family: var(--display);
  font-size: 1.55rem;
  font-weight: 400;
}

.process-step p {
  max-width: 38rem;
  margin-bottom: 0;
  color: var(--muted);
}

.lab-section {
  background: var(--ice-light);
}

.scenario-shell {
  display: grid;
  grid-template-columns: 0.82fr 1.18fr;
  gap: clamp(2rem, 6vw, 7rem);
  margin-top: 4rem;
  padding: clamp(1.5rem, 4vw, 4rem);
  border: 1px solid rgba(33,28,27,0.22);
  background: rgba(250,247,241,0.52);
}

.scenario-controls {
  display: flex;
  flex-direction: column;
  gap: 2.1rem;
}

.control-header {
  display: flex;
  align-items: baseline;
  justify-content: space-between;
  gap: 1rem;
  margin-bottom: 0.75rem;
}

.control-header label {
  font-size: 0.7rem;
  font-weight: 700;
  letter-spacing: 0.09em;
  text-transform: uppercase;
}

.control-header output {
  color: var(--plum);
  font-family: var(--display);
  font-size: 1.35rem;
}

input[type="range"] {
  width: 100%;
  height: 2px;
  appearance: none;
  border-radius: 99px;
  background: rgba(33,28,27,0.25);
}

input[type="range"]::-webkit-slider-thumb {
  width: 1.15rem;
  height: 1.15rem;
  appearance: none;
  border: 3px solid var(--ice-light);
  border-radius: 50%;
  background: var(--plum);
  box-shadow: 0 0 0 1px var(--plum);
  cursor: pointer;
}

input[type="range"]::-moz-range-thumb {
  width: 0.9rem;
  height: 0.9rem;
  border: 3px solid var(--ice-light);
  border-radius: 50%;
  background: var(--plum);
  box-shadow: 0 0 0 1px var(--plum);
  cursor: pointer;
}

.scenario-output {
  padding-left: clamp(1rem, 4vw, 4rem);
  border-left: 1px solid rgba(33,28,27,0.2);
}

.scenario-output > p {
  color: var(--muted);
  font-size: 0.76rem;
  font-weight: 700;
  letter-spacing: 0.09em;
  text-transform: uppercase;
}

.metric {
  display: grid;
  grid-template-columns: 1fr auto;
  gap: 0.6rem 2rem;
  align-items: end;
  padding: 1.2rem 0;
  border-bottom: 1px solid rgba(33,28,27,0.18);
}

.metric-label {
  font-family: var(--display);
  font-size: clamp(1.25rem, 2vw, 1.7rem);
}

.metric-value {
  color: var(--plum);
  font-family: var(--display);
  font-size: clamp(1.6rem, 3vw, 2.5rem);
}

.metric-bar {
  grid-column: 1 / -1;
  height: 0.25rem;
  overflow: hidden;
  background: rgba(33,28,27,0.12);
}

.metric-bar span {
  display: block;
  width: 50%;
  height: 100%;
  background: var(--plum);
  transition: width 300ms ease;
}

.scenario-disclaimer {
  margin: 1.2rem 0 0;
  color: var(--muted);
  font-size: 0.72rem;
  line-height: 1.5;
}

.principles-grid {
  display: grid;
  grid-template-columns: repeat(3, 1fr);
  gap: 1px;
  background: var(--line);
  border: 1px solid var(--line);
}

.principle {
  min-height: 20rem;
  padding: clamp(1.5rem, 3vw, 3rem);
  background: var(--paper);
}

.principle span {
  display: block;
  margin-bottom: 5rem;
  color: var(--plum);
  font-family: var(--display);
}

.principle h3 {
  margin-bottom: 1rem;
  font-family: var(--display);
  font-size: 1.75rem;
  font-weight: 400;
}

.principle p {
  margin-bottom: 0;
  color: var(--muted);
}

.site-footer {
  padding-top: clamp(4rem, 8vw, 7rem);
  padding-bottom: 2rem;
  background: var(--ink);
  color: white;
}

.footer-grid {
  display: grid;
  grid-template-columns: 1.2fr 0.8fr;
  gap: clamp(3rem, 8vw, 8rem);
  align-items: end;
  padding-bottom: 5rem;
}

.footer-brand {
  max-width: 9em;
  margin-bottom: 1.8rem;
  font-family: var(--display);
  font-size: clamp(2.8rem, 5vw, 5rem);
  font-weight: 400;
  letter-spacing: -0.04em;
  line-height: 0.95;
}

.footer-note {
  max-width: 34rem;
  color: rgba(255,255,255,0.6);
}

.footer-links {
  display: grid;
  grid-template-columns: repeat(2, 1fr);
  gap: 0.8rem 2rem;
}

.footer-links a {
  padding: 0.6rem 0;
  border-bottom: 1px solid rgba(255,255,255,0.2);
  font-size: 0.73rem;
  font-weight: 700;
  letter-spacing: 0.09em;
  text-transform: uppercase;
}

.footer-links a:hover {
  color: var(--ice);
}

.footer-bottom {
  display: flex;
  align-items: start;
  justify-content: space-between;
  gap: 2rem;
  padding-top: 1.4rem;
  border-top: 1px solid rgba(255,255,255,0.16);
  color: rgba(255,255,255,0.5);
  font-size: 0.68rem;
  line-height: 1.5;
}

.reveal {
  opacity: 0;
  transform: translateY(2rem);
  transition: opacity 700ms cubic-bezier(.2,.7,.2,1), transform 700ms cubic-bezier(.2,.7,.2,1);
}

.reveal.is-visible {
  opacity: 1;
  transform: translateY(0);
}

@media (prefers-reduced-motion: reduce) {
  html { scroll-behavior: auto; }
  *, *::before, *::after {
    animation-duration: 0.01ms !important;
    animation-iteration-count: 1 !important;
    transition-duration: 0.01ms !important;
  }
  .reveal { opacity: 1; transform: none; }
}

@media (max-width: 1000px) {
  .home-hero-grid,
  .page-hero-grid,
  .mission-grid,
  .grant-band,
  .model-intro,
  .scenario-shell {
    grid-template-columns: 1fr;
  }

  .home-hero-image-wrap {
    min-height: 35rem;
    margin-top: 3rem;
  }

  .home-hero-copy .hero-line:nth-child(2) {
    transform: translateX(0);
  }

  .hero-intro {
    grid-template-columns: 1fr;
  }

  .mission-grid {
    gap: 5rem;
  }


  .page-hero-aside {
    max-width: 42rem;
  }

  .team-lead {
    grid-template-columns: 1fr;
  }

  .pi-card {
    min-height: 24rem;
  }

  .pi-monogram {
    font-size: 15rem;
  }

  .model-intro h2 {
    position: static;
  }

  .scenario-output {
    padding-top: 2rem;
    padding-left: 0;
    border-top: 1px solid rgba(33,28,27,0.2);
    border-left: 0;
  }
}

@media (max-width: 760px) {
  :root {
    --radius-lg: 0 4rem 0 4rem;
  }

  .nav-shell {
    min-height: 4.8rem;
  }

  .menu-toggle {
    display: block;
  }

  .nav-links {
    position: fixed;
    inset: 4.8rem 0 0;
    display: flex;
    flex-direction: column;
    align-items: flex-start;
    gap: 0;
    padding: 2rem var(--pad) 4rem;
    background: var(--paper);
    transform: translateX(100%);
    transition: transform 260ms ease;
  }

  .menu-open .nav-links {
    transform: translateX(0);
  }

  .nav-link {
    width: 100%;
    padding: 1.15rem 0;
    border-bottom: 1px solid var(--line);
    font-family: var(--display);
    font-size: 2rem;
    font-weight: 400;
    letter-spacing: -0.02em;
    text-transform: none;
  }

  .home-hero {
    min-height: auto;
  }

  .home-hero-grid {
    display: block;
  }

  .home-hero-copy {
    padding-top: 2rem;
  }

  .home-hero-image-wrap {
    min-height: 30rem;
  }

  .stat-row,
  .pillar-grid,
  .team-grid,
  .principles-grid,
  .footer-grid {
    grid-template-columns: 1fr;
  }

  .stat-row {
    gap: 1px;
  }

  .stat {
    min-height: auto;
  }

  .pillar-grid {
    border-top: 1px solid rgba(255,255,255,0.2);
  }

  .pillar,
  .pillar + .pillar {
    min-height: auto;
    padding: 2rem 0 3rem;
    border-right: 0;
    border-bottom: 1px solid rgba(255,255,255,0.2);
  }

  .pillar-number {
    margin-bottom: 4rem;
  }

  .section-heading-row {
    display: block;
  }

  .section-heading-row .button {
    margin-top: 2rem;
  }

  .team-member,
  .team-member:nth-child(even) {
    padding: 2rem 0;
    border-left: 0;
  }

  .publication {
    grid-template-columns: 4rem minmax(0, 1fr);
  }

  .pub-arrow {
    display: none;
  }

  .comet-hero {
    min-height: 44rem;
  }

  .comet-hero-image {
    object-position: 37% center;
  }

  .process-step {
    grid-template-columns: 2.7rem 1fr;
  }

  .principle {
    min-height: auto;
  }

  .principle span {
    margin-bottom: 3rem;
  }

  .footer-grid {
    gap: 4rem;
  }

  .footer-bottom {
    display: block;
  }
}

@media (max-width: 480px) {
  .brand {
    min-width: 0;
    max-width: 11rem;
    font-size: 0.63rem;
  }

  .display-xl {
    font-size: clamp(3.55rem, 18vw, 5rem);
  }

  .page-hero {
    padding-top: 3.6rem;
  }

  .home-hero-image-wrap {
    min-height: 25rem;
  }


  .grant-fact {
    grid-template-columns: 1fr;
    gap: 0.35rem;
  }

  .team-member {
    grid-template-columns: minmax(0, 1fr);
  }

  .publication {
    grid-template-columns: 1fr;
    gap: 0.6rem;
  }

  .scenario-shell {
    padding: 1.25rem;
  }

  .footer-links {
    grid-template-columns: 1fr;
  }
}

/* COMET-Lung Online landing page */
.comet-launch {
  display: inline-flex;
  align-items: center;
  gap: 1.2rem;
  max-width: 100%;
  padding: 1.2rem 1.5rem;
  border: 1px solid var(--ice);
  background: var(--ice);
  color: var(--ink);
  transition: background 180ms ease, transform 180ms ease;
}
.comet-launch[href]:hover { background: white; transform: translateY(-3px); }
.comet-launch:focus-visible { outline: 3px solid white; outline-offset: 6px; }
.comet-launch[aria-disabled="true"] { cursor: default; }
.comet-launch-icon { flex: 0 0 3.5rem; width: 3.5rem; height: 3.5rem; }
.comet-launch-icon svg { width: 100%; height: 100%; }
.comet-launch strong { display: block; font-size: 1.1rem; }
.comet-launch-detail { display: block; margin-top: 0.2rem; font-size: 0.85rem; }
.comet-launch-arrow { margin-left: 1rem; font-size: 1.8rem; }
.launch-status { margin: 0.85rem 0 1.7rem; color: rgba(255,255,255,0.8); font-size: 0.85rem; }
.comet-overview-link { font-size: 0.8rem; border-bottom: 1px solid rgba(255,255,255,0.5); padding-bottom: 0.4rem; }
.comet-content { display: grid; grid-template-columns: minmax(0, 1fr) minmax(16rem, 0.43fr); gap: clamp(3rem, 7vw, 7rem); scroll-margin-top: 7rem; }
.comet-copy section + section { margin-top: 3.8rem; padding-top: 3.8rem; border-top: 1px solid var(--line); }
.comet-copy h2 { font-family: var(--display); font-size: clamp(2rem, 3vw, 2.8rem); font-weight: 400; line-height: 1.15; letter-spacing: -0.035em; }
.comet-copy p, .comet-outcomes { font-size: 1.05rem; line-height: 1.8; }
.comet-copy p a { text-decoration: underline; text-underline-offset: 3px; color: var(--plum); }
.comet-outcomes { padding-left: 1.25rem; margin: 1.6rem 0; }
.comet-outcomes li { padding-left: 0.4rem; margin-bottom: 0.55rem; }
.comet-outcomes li::marker { color: var(--plum); }
.comet-reading { align-self: start; padding: 2rem; background: var(--paper-light); border-top: 3px solid var(--plum); }
.comet-reading h2 { font-family: var(--display); font-size: 1.8rem; font-weight: 400; line-height: 1.15; }
.comet-paper { display: block; padding: 1.6rem 0; border-top: 1px solid var(--line); }
.comet-paper span { display: block; color: var(--plum); font-size: 0.68rem; font-weight: 700; letter-spacing: 0.07em; text-transform: uppercase; }
.comet-paper h3 { margin: 0.8rem 0; font-family: var(--display); font-size: 1.3rem; font-weight: 400; line-height: 1.3; }
.comet-paper p { color: var(--muted); font-size: 0.8rem; }
.comet-paper:hover h3 { text-decoration: underline; text-underline-offset: 3px; }
.comet-reading > .text-link { margin-top: 1rem; }
@media (max-width: 900px) {
  .comet-content { grid-template-columns: 1fr; }
}
@media (max-width: 480px) {
  .comet-launch { gap: 0.8rem; padding: 1rem; }
  .comet-launch-icon { flex-basis: 2.7rem; width: 2.7rem; height: 2.7rem; }
  .comet-launch strong { font-size: 0.92rem; }
  .comet-launch-arrow { margin-left: 0; }
  .comet-reading { padding: 1.5rem; }
}

/* The watercolor layers begin together and gently fan toward the right. */
.home-hero-grid {
  grid-template-columns: minmax(0, 0.85fr) minmax(0, 1.15fr);
  align-items: center;
}
.lung-art {
  position: relative;
  width: 100%;
  aspect-ratio: 1.4;
  isolation: isolate;
}
.lung-layer {
  position: absolute;
  top: 8%;
  left: 0;
  width: 64%;
  height: 84%;
  object-fit: contain;
  mix-blend-mode: multiply;
  transform: translateX(var(--lung-shift));
  animation: lung-unfold 3.8s cubic-bezier(0.22, 0.61, 0.36, 1) 0.4s both;
}
.lung-layer-back {
  --lung-shift: 0%;
  opacity: 0.22;
  filter: grayscale(1) sepia(0.4) hue-rotate(155deg) saturate(1.8);
}
.lung-layer-middle { --lung-shift: 28%; opacity: 0.26; }
.lung-layer-front { --lung-shift: 56%; opacity: 0.62; }
@keyframes lung-unfold {
  from { transform: translateX(0); }
  to { transform: translateX(var(--lung-shift)); }
}
@media (max-width: 1000px) {
  .home-hero-grid { grid-template-columns: 1fr; }
  .lung-art { max-width: 48rem; margin: 1rem auto 0; }
}
@media (prefers-reduced-motion: reduce) {
  .lung-layer { animation: none; }
}
.home-hero-grid > .hero-intro { grid-column: 1 / -1; }

/* Native disclosure keeps citations and abstracts usable without JavaScript. */
details.publication {
  display: block;
  padding: 0;
}
.publication-summary {
  display: grid;
  grid-template-columns: 6rem minmax(0, 1fr) 2.5rem;
  gap: clamp(1.2rem, 3vw, 3rem);
  align-items: start;
  padding: clamp(1.8rem, 3vw, 3rem) 0;
  cursor: pointer;
  list-style: none;
}
.publication-summary::-webkit-details-marker { display: none; }
.publication-summary::marker { content: ""; }
.publication-summary:focus-visible {
  outline: 2px solid var(--plum);
  outline-offset: 5px;
}
.publication-summary:hover h2 { color: var(--plum); }
.publication h2 { margin-bottom: 0.8rem; }
.pub-contribution {
  max-width: 66ch;
  margin-bottom: 1.2rem;
  color: var(--ink);
  font-size: 1rem;
  line-height: 1.6;
}
.pub-expand {
  display: grid;
  width: 2.3rem;
  height: 2.3rem;
  place-items: center;
  border: 1px solid var(--line);
  border-radius: 50%;
  color: var(--plum);
  font-size: 1.5rem;
  line-height: 1;
}
.pub-expand::before { content: "+"; }
.publication[open] .pub-expand::before { content: "−"; }
.publication-expanded {
  margin-left: calc(6rem + clamp(1.2rem, 3vw, 3rem));
  margin-right: calc(2.5rem + clamp(1.2rem, 3vw, 3rem));
  padding: 0 0 2.5rem;
}
.publication-expanded h3 {
  margin-bottom: 0.25rem;
  font-family: var(--display);
  font-size: 1.6rem;
  font-weight: 400;
}
.pub-abstract-source { color: var(--muted); font-size: 0.75rem; margin-bottom: 1.5rem; }
.pub-abstract { max-width: 78ch; font-size: 0.95rem; line-height: 1.8; }
.pub-abstract p { margin-bottom: 1.1rem; }
.pub-article-link { display: inline-block; margin-top: 0.8rem; }
.sr-only {
  position: absolute;
  width: 1px;
  height: 1px;
  padding: 0;
  margin: -1px;
  overflow: hidden;
  clip-path: inset(50%);
  white-space: nowrap;
  border: 0;
}
@media (max-width: 760px) {
  .publication-summary { grid-template-columns: 4rem minmax(0, 1fr) 2.3rem; gap: 1rem; }
  .publication-expanded { margin-left: 5rem; margin-right: 0; }
}
@media (max-width: 480px) {
  .publication-summary { grid-template-columns: minmax(0, 1fr) 2.3rem; }
  .publication-summary .pub-year { grid-column: 1; }
  .publication-summary > div { grid-column: 1 / -1; grid-row: 2; }
  .publication-summary .pub-expand { grid-column: 2; grid-row: 1; }
  .publication-expanded { margin-left: 0; }
}

.team-credentials {
  display: block;
  margin-top: 0.55rem;
  font-family: var(--sans);
  font-size: 1rem;
  font-weight: 400;
  line-height: 1.4;
  letter-spacing: normal;
}

/* Source code and repository announcements */
.repository-card {
  display: grid;
  grid-template-columns: minmax(0, 1.4fr) minmax(16rem, .8fr);
  gap: clamp(2rem, 6vw, 6rem);
  border-top: 1px solid var(--line);
  padding-top: 3rem;
}
.repository-card p { max-width: 65ch; }
.repository-actions { display: flex; flex-direction: column; align-items: flex-start; justify-content: center; gap: 1.5rem; }
.source-news { background: var(--paper-light); }
.source-news .lead { max-width: 52ch; }
.source-updates { margin-top: 3rem; border-top: 1px solid var(--line); }
.source-updates-empty { padding-top: 1.5rem; color: var(--muted); }
.source-update { display: grid; grid-template-columns: 10rem minmax(0, 1fr); gap: 2rem; padding: 2rem 0; border-bottom: 1px solid var(--line); }
.source-update time { color: var(--muted); }
.source-update p { max-width: 65ch; }
@media (max-width: 760px) {
  .repository-card, .source-update { grid-template-columns: 1fr; }
}

/* Keep the expanded banner and five navigation links comfortable on tablets. */
@media (min-width: 761px) and (max-width: 1150px) {
  .menu-toggle { display: block; }
  .nav-links {
    position: absolute;
    inset: 100% 0 auto;
    flex-direction: column;
    align-items: stretch;
    gap: 0;
    padding: 1rem var(--pad) 2rem;
    background: var(--paper);
    visibility: hidden;
  }
  .menu-open .nav-links { visibility: visible; }
  .nav-link { padding: 1rem 0; border-bottom: 1px solid var(--line); }
}